Harman Patil (Editor)

London Clay

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Type
  
Formation

Region
  
southern England

Unit of
  
Thickness
  
up to 150 m

Overlies
  
Harwich Formation

London Clay httpsc2staticflickrcom434903776171422615f

Underlies
  

A new fossil bramoides from the eocene london clay re aligned with the modern genus gasterochisma


The London Clay Formation is a marine geological formation of Ypresian (Lower Eocene Epoch, c. 56–49 Ma) age which crops out in the southeast of England. The London Clay is well known for its fossil content. The fossils from the Lower Eocene indicate a moderately warm climate, the tropical or subtropical flora. Though sea levels changed during the deposition of the Clay, the habitat was generally a lush forest – perhaps like in Indonesia or East Africa today – bordering a warm, shallow ocean.

Contents

The London Clay is a stiff bluish clay which becomes brown when weathered. Nodular lumps of pyrite and crystals of selenite (sometimes called "waterstones") frequently exist in the clay, and large septarian concretions are also common. These have been used in the past for making cement. They were once dug for this purpose at Sheppey, near Sittingbourne, and at Harwich, and also dredged, off the Hampshire coast. The clay is still used commercially for making bricks, tiles, and coarse pottery in places such as Michelmersh in Hampshire. It is infertile for gardens and crops.

London Clay Victorian Highbury London Clay amp the Brickfields Highbury

Distribution and geology

London Clay London Clay Wikipedia

The London Clay is well developed in the London Basin, where it thins westwards from around 150 metres (492 feet) in Essex and north Kent to around 4.6 metres (15 feet) in Wiltshire. though it is not frequently exposed as it is to a great extent covered by more recent Neogene sediments and Pleistocene gravel deposits. One location of particular interest is Oxshott Heath, where the overlying sand and the London Clay layers are exposed as a sand escarpment, rising approximately 25 metres (82 feet). This supported a thriving brick industry in the area until the 1960s. The London Clay is also well developed in the Hampshire Basin, where an exposure 91 metres (299 ft) thick occurs at Whitecliff Bay on the Isle of Wight and around 101 metres (331 ft) is spread along 6 kilometres (4 miles) of foreshore at Bognor Regis, West Sussex.

The clay was deposited in a sea up to 200 metres (660 ft) deep at the eastern end. Up to five cycles of deposition (representing transgression followed by shallowing of the sea) have been found, most markedly at the shallower, western end. Each cycle begins with coarser material (sometimes including rounded flint pebbles), followed by clay which becomes increasingly sandy. The final cycle ends with the Claygate Beds.

Claygate Beds

London Clay Modelling Volume Change Potential in the London Clay

The youngest part of the London Clay, known as the Claygate Beds or Claygate Member forms a transition between the clay and the sandier Bagshot Beds above. This is shown separately on many geological maps, and often caps hills. It is up to 15 metres (49 ft) thick at Claygate, Surrey. It is now believed to be diachronous, with the formation at Claygate for example being the same age as the end of the fourth cycle of deposition further east.

Engineering

The presence of a thick layer of London Clay underneath London itself, providing a soft yet stable environment for tunnelling, was instrumental in the early development of the London Underground, although this is also the reason why London has no true skyscraper buildings, at least to the same degree as many other cities throughout the world. Erecting tall buildings in London requires very deep, large and costly piled foundations.

London Clay is highly susceptible to volumetric changes depending upon its moisture content. During exceptionally dry periods or where the moisture is extracted by tree root activity, the clay can become desiccated and shrink in volume, and conversely swell again when the moisture content is restored. This can lead to many problems near the ground surface, including structural movement and fracturing of buildings, fractured sewers and service pipes/ducts and uneven and damaged road surfaces and pavings. Such damage is recognised to be covered by the interpretation of subsidence in buildings insurance policies, and the periods of dry weather in 1976/77 and 1988/92, in particular, led to a host of insurance claims. As a result, many insurance companies have now increased the cost of premiums for buildings located in the most susceptible areas where damage occurred, where the clay is close to the surface.

Tunnels in London Clay

The London Clay is an ideal medium for driving tunnels, which is why the London Tube railway network expanded quickly north of the Thames, but south of the Thames the stratum at tube level is water-bearing sand and gravel (not good for tunnelling) with London Clay below, which is why there are few tube tunnels there. London Clay has a stand-up time long enough to enable support to be installed without urgency. It is also almost waterproof, resulting in virtually no seepage of groundwater into the tunnel. It is over-consolidated, which means that it is under pressure, and expands upon excavation, thus gradually loading the support, i.e. it is not necessary to stress the support against the ground.

Agriculture

Due to its impermeability especially when exposed by ploughing, London clay does not make good agricultural soil. In Middlesex, it has historically been called "ploughing up poison."

Fossil flora and fauna

Notable coastal exposures from which fossils can be collected are on the Isle of Sheppey in Kent and Walton-on-the-Naze, Essex in the London Basin, and Bognor Regis in the Hampshire Basin.

Plant fossils, especially seeds and fruits, are found in abundance and have been collected from the London Clay for almost 300 years. Some 350 named species of plant have been found, making the London Clay flora one of the world's most diverse for fossil seeds and fruits. The flora includes plant types found today in tropical forests of Asia and demonstrates the much warmer climate of the Eocene epoch, with plants such as Nypa (Nipah palms) and other palms being frequently encountered. The following plants list is incomplete and is based on the research by Marjorie Chandler. and research works done by the paleobotanist Steven R. Manchester and by professor of plant palaeobiology Margaret Collinson.

Pteridophytes

  • Pteridaceae
  • Acrostichum – mangrove fern
  • Salviniaceae – water ferns
  • Azolla
  • Gymnosperms

  • Araucariaceae – monkey puzzle, bunya pine, and Norfolk pine
  • Araucarites spp.
  • Doliostrobus stenbergi - fossils in the past named †Araucarites are said to be of this genus, it is now placed in its own family, †Doliostrobaceae.
  • Pinaceae
  • Pinus spp.
  • Taxaceae
  • ?Taxaceae indet.
  • Cephalotaxus bowerbanki
  • Cupressaceae
  • Cupressinites spp.
  • Libocedrus adpressa
  • Quasisequoia sp.
  • Sequoia couttsiae
  • Podocarpaceae
  • Podocarpus argillaelondinensis?
  • Angiosperms

  • Arecaceae – palms
  • Caryotispermum cantiense
  • Livistona minima
  • Nypa burtini (syn. Nipa burtini)
  • Oncosperma anglica
  • Palmospermum bracknellense
  • Palmospermum cooperi
  • Palmospermum davisi
  • Palmospermum elegans
  • Palmospermum jenkinsi
  • Palmospermum minutum
  • Palmospermum ornatum
  • Palmospermum ovale
  • Palmospermum subglobulare
  • Sabal grandisperma
  • Cyclanthaceae
  • Cyclanthus lakensis
  • Cyperaceae - sedges
  • Caricoidea obovata
  • Polycarpella caespitosa
  • Posidoniaceae - seagrass family
  • Posidonia parisiensis
  • Nymphaeaceae – water lilies
  • Protobarclaya eocenica
  • Magnoliaceae
  • Liriodendron gardneri - tulip tree
  • Magnolia angusta
  • Magnolia crassa
  • Magnolia davisi
  • Magnolia lata
  • Magnolia lobata
  • Magnolia oblonga
  • Magnolia pygmaea
  • Magnolia rugosa
  • Magnolia subquadrangularis
  • Magnolia symmetrica
  • Myristicaceae - nutmeg family
  • Myristicacarpum chandlerae
  • Annonaceae - the custard apple family
  • Anonaspermum anonijorme - related to the genera Anonna and Polyalthia, most similar species is Annona muricata
  • Anonaspermum cerebellatum - Uvaria, Melodorum and Orophea shows closest resemblance to the fossil
  • Anonaspermum commune
  • Anonaspermum complanatum - related to Anonna and Melodorum
  • Anonaspermum corrugatum - related to the genera Anonna and Melodorum
  • Anonaspermum minimum - related to Dasymaschalon clusiflorum
  • Anonaspermum obscurum
  • Anonaspermum ovale - related to the genera Anonna and Melodorum
  • Anonaspermum pulchrum - related to Dasymaschalon clusiflorum
  • Anonaspermum punctatum - related to the genera Bocagea, Orophea, Unonopsis, and Guatteria
  • Anonaspermum rotundatum - distantly related to Polyauhia
  • Anonaspermum rugosum
  • Anonaspermum subcompressum
  • Lauraceae – avocado and cinnamon family
  • Beilschmiedia bognorensis
  • Beilschmiedia eocenica
  • Beilschmiedia gigantea
  • Beilschmiedia oviformis
  • Beilschmiedia pyriformis
  • Cinnamomum grande
  • Cinnamomum globulare
  • Cinnamomum oblongum
  • Cinnamomum ovoideum
  • Crowella globosa
  • Endiandra spp.
  • Laurocalyx bowerbanki
  • Laurocalyx dubius
  • Laurocalyx fibrotorulosus
  • Laurocalyx globularis
  • Laurocalyx magnus
  • Laurocarpum crassum
  • Laurocarpum cupuliferum
  • Laurocarpum davisi
  • Laurocarpum inornatum
  • Laurocarpum minimum
  • Laurocarpum minutissimum
  • Laurocarpum ovoideum
  • Laurocarpum paradoxum
  • Laurocarpum proteum
  • Laurocarpum pyrocarpum
  • Laurocarpum sheppeyense
  • Litsea pyriformis
  • Protoaltingia europaea
  • Protoravensara sheppeyensis
  • Dilleniaceae
  • Hibbertia bognorensis
  • Tetracera ?cantiensis
  • Tetracera crofti
  • Tetracera eocenica
  • Platanaceae – sycamore or plane tree
  • Plataninium decipiens
  • Proteaceae
  • Dryandra acutiloba
  • Betulaceae – birch, alder and hornbeams
  • Alnus richardsoni (syn. Petrophiloides richardsoni) – an alder
  • Hamamelidaceae – witch-hazel family
  • Corylopsis? bognorensis
  • Corylopsis? latisperma
  • Corylopsis venablesi
  • Protoaltingia europaea
  • Myricaceae - wax-myrtle, bayberry family
  • Myrica boveyana
  • Juglandaceae – walnut, hickory and pecan family
  • Juglandicarya bognorensis
  • Juglandicarya cooperi
  • Juglandicarya depressa - the most common Juglandacarya species in the London Clay
  • Juglandicarya lubbocki
  • Juglandicarya minuta
  • Petrophiloides richardsoni - among the two most abundant Juglandaceae species in the London Clay
  • Pterocaryopsis bognorensis - related to the genus Pterocarya
  • Trochodendraceae
  • Trochodendron pauciseminum
  • Sabiaceae
  • Bognoria venablesi
  • Meliosma cantiensis
  • Meliosma jenkinsi
  • Meliosma sheppeyensis
  • Menispermaceae – moon seed family
  • Bowerbankella tiliacoroidea
  • Davisicarpum gibbosum
  • Eohypserpa parsonsi
  • Frintonia ornata
  • Menispermicarpum rariforme
  • Menispermoxylon - close to the extant genus Tinomiscium
  • Microtinomiscium foveolatum
  • Palaeococculus lakensis
  • Palaeosinomenium pulchrum
  • Tinomiscoidea scaphiformis
  • Tinospora excavata
  • Wardenia davisi
  • Aristolochiaceae - birthwort family
  • Aristolochia sp.
  • Torricelliaceae
  • Toricellia sp. - very similar to Torricellia bonesii from the Clarno Formation of Oregon
  • Altingiaceae
  • Liquidambar palaeocenica - sweetgum
  • Oleaceae
  • Fraxinus sp.
  • Styracaceae
  • Styrax sp.
  • Symplocaceae – sapphire-berry, sweet leaf.
  • Symplocos curvata
  • Symplocos quadrilocularis
  • Symplocos trilocularis
  • Symplocos bognorensis
  • Ericaceae
  • Leucopogon quadrilocularis
  • Pentaphylacaceae
  • Sapotaceae
  • Sapoticarpum rotundatum
  • Sapoticarpum latum
  • Sapoticarpum duhium
  • Sapotispermum sheppeyense - allied to Chrysophyllum and Sideroxylon
  • Myrtaceae
  • Hightea elliptica
  • Hightea turgida
  • Myrtospermum variabile
  • Palaeorhodomyrtus subangulata - allied to Rhodomyrtus
  • Fabaceae
  • Dalbergia sp.
  • Leguminocarpon nervosum
  • Mimosites browniana
  • Myrsinaceae
  • Ardisia eocenica
  • Rhizophoraceae - mangrove
  • Palaeobruguiera elongata
  • Palaeobruguiera lata
  • Malpighiaceae
  • Salicaceae
  • Oncoba variabilis
  • Saxifragispermum spinosissimum
  • Linaceae
  • Decaplatyspermum bowerbanki
  • Wetherellia variahilis - related to Hugonia
  • Nyctaginaceae - the four o'clock family
  • Pisonia sp. - the birdcatcher tree
  • Olacaceae
  • Erythropalum europaeum - Only one extant species of this genus which is a scandent shrubs or liana
  • Erythropalum jenkinsi
  • Erythropalum turbinatum
  • Olax depressa
  • Boraginaceae
  • Ehretia clausentia
  • Ehretia ehretioides
  • Solanaceae
  • Cantisolanum daturoides
  • Apocynaceae - the dogbane family
  • Ochrosella ovalis
  • Ochrosoidea sheppeyensis
  • Burseraceae - the incense tree family
  • Bursericarpum aldwickense
  • Bursericarpum bognorense
  • Bursericarpum ovale
  • Bursericarpum venablesi
  • Palaeobursera bognorensis
  • Protocommiphora europaea
  • Anacardiaceae
  • Choerospondias sheppeyensis
  • Dracontomelon minimum
  • Dracontomelon subglobosum
  • Lannea europaea
  • Lannea jenkinsi
  • Lobaticarpum variabile
  • Odina europaea
  • Odina jenkinsi
  • Odina subreniformis
  • Pseudosclerocarya subalata
  • Spondiaecarpon operculatum - according to several botanists, the pyritized specimens, originally described as Spondiaecarpon operculatum, represent locule casts of Torricellia sp.
  • Onagraceae
  • Palaeeucharidium cellulare - allied to Eucharidium
  • Lythraceae
  • Cranmeria trilocularis
  • Minsterocarpum alatum - closely related to the crape myrtle or crepe myrtle genus Lagerstroemia
  • Pachyspermum quinqueloculare
  • Tamesicarpum polyspermum
  • Malvaceae
  • Cantitilia polysperma
  • Elaeocarpaceae
  • Echinocarpus sheppeyensis
  • Moraceae – mulberry and fig family
  • ?Morus sp.
  • Urticaceae
  • Urticicarpum scutellum
  • Euphorbiaceae - the spurge family
  • Euphorbiospermum bognorense
  • Euphorbiospermum cooperi
  • Euphorbiospermum eocenicum
  • Euphorbiospermum obliquum
  • Euphorbiospermum subglobulare
  • Euphorbiospermum subovoideum
  • Euphorbiospermum venablesi
  • Euphorbiotheca minima
  • Euphorbiotheca sheppeyensis
  • Lagenoidea bilocularis
  • Lagenoidea trilocularis
  • Wetherellia variabilis
  • Cucurbitaceae - the gourd family
  • Cucurbitospermum cooperi
  • Cucurbitospermum equiaelaterale
  • Cucurbitospermum sheppeyense
  • Cucurbitospermum triangulare
  • Vitaceae
  • Tetrastigma corrugata
  • Tetrastigma davisi
  • Tetrastigma elliotti
  • Tetrastigma sheppeyensis
  • Vitacexoylon sp. - close to the extant genus Rhoicissus
  • Vitis bilobata
  • Vitis bognorensis
  • Vitis bracknellensis
  • Vitis elegans
  • Vitis longisulcata
  • Vitis magnisperma
  • Vitis obovoidea
  • Vitis platyformis
  • Vitis pygmaea
  • Vitis rectisulcata
  • Vitis semenlabruscoides
  • Vitis subglobosa
  • Vitis venablesi
  • Sapindaceae - soapberry
  • Cupanoides grandis
  • Cupanoides tumidus
  • Palaeallophylus minimus
  • Palaeallophyllus ovoideus
  • Palaeallophylus rotundatus
  • Palaealectryon spirale
  • Sapindospermum cooperi
  • Sapindospermum davisi
  • Sapindospermum grande
  • Sapindospermum jenkinsi
  • Sapindospermum ovoideum
  • Sapindospermum revolutum
  • Sapindospermum subovatum
  • Meliaceae - the mahogany family
  • Cedrela sp.
  • Melicarya variabili
  • Toona sulcata
  • Rutaceae - the rue family
  • Canticarya gracilis
  • Canticarya ovalis
  • Canticarya sheppeyensis
  • Canticarya ventricosa
  • Caxtonia elongata
  • Caxtonia glandulosa
  • Caxtonia rutacaeformis
  • Citrispermum sheppeyense
  • Clausenispermum dubium
  • Eozanthoxylon glandulosum
  • Euodia costata
  • Rutaspermum bognorense
  • Rutaspermum minimum
  • Shrubsolea jenkinsi
  • Zanthoxylon compression
  • Zanthoxylon bognorense
  • Celastraceae - the staff vine or bittersweet family
  • Canticarpum celastroides
  • Cathispermum pulchrum
  • Celastrinoxylon ramunculiformis
  • Sterculiaceae
  • Sphinxia ovalis
  • Cornaceae
  • Beckettia mastixioides
  • Cornus ettingshausenii
  • Dunstania ettinghauseni
  • Dunstania multilocularis
  • Langtonia bisulcata
  • Lanfrancia subglobosa
  • Mastixia cantiensis
  • Mastixia grandis
  • Mastixia parva
  • Portnallia bognorensis
  • Portnallia sheppeyensis
  • Nyssaceae - the tupelo family
  • Nyssa bilocularis
  • Nyssa cooperi
  • Palaeonyssa multilocularis
  • Curtisiaceae
  • Curtisia quadrilocularis
  • Alangiaceae
  • Alangium jenkinsi
  • Icacinaceae
  • Faboidea crassicutis
  • Icacinicarya amygadaloidea
  • Icacinicarya bartonensis
  • Icacinicarya becktonensis
  • Icacinicarya bognorensis
  • Icacinicarya echinata
  • Icacinicarya elegans
  • Icacinicarya emarginata
  • Icacinicarya forbesii
  • Icacinicarya foveolata
  • Icacinicarya glabra
  • Icacinicarya inornata
  • Icacinicarya jenkinsi
  • Icacinicarya minima
  • Icacinicarya mucronata
  • Icacinicarya nodulifera
  • Icacinicarya ovalis
  • Icacinicarya ovoidea
  • Icacinicarya platycarpa
  • Icacinicarya pygmaea
  • Icacinicarya reticulata
  • Icacinicarya rotundata
  • Icacinicarya transversalis
  • Iodes acutiform
  • Iodes bilinica
  • Iodes corniculata
  • Iodes davisii
  • Iodes eocenica
  • Iodes multireticulata
  • Natsiatum eocenicum
  • Palaeophytocrene ambigua
  • Palaeophytocrene foveolata
  • Sphaeriodes ventricosa
  • Stizocarya communis
  • Animals

    Animal fossils include bivalves, gastropods, nautilus, worm tubes, brittle stars and starfish, crabs, lobsters, fish (including shark and ray teeth), reptiles (particularly turtles), and a large diversity of birds. A few mammal remains have also been recorded. Preservation varies; articulated skeletons are generally rare. Of fish, isolated teeth are very frequent. Bird bones are not infrequently encountered compared to other lagerstätten, but usually occur as single bones and are often broken.

    The following fauna species list follows Clouter (2007).

    Vertebrates

    Mammals
  • Argillotherium
  • Coryphodon eocaenus – a pantodont
  • Hyracotherium – the famous horse ancestor
  • Oxyaena – a creodont
  • Platychoerops richardsoni - from Herne Bay, a primate belonging to the order Plesiadapiformes
  • Birds
  • Anatalavis oxfordi – a waterbird possibly related to the magpie-goose of Australia
  • Argillipes – perhaps a landfowl
  • Dasornis, Odontopteryx and Pseudodontornis – pseudo-tooth birds
  • Eocolius – a coliiform
  • Eostrix – an owl
  • Gastornis - from the Isle of Grain, a very large flightless bird
  • Halcyornis – a parrot or roller relative
  • Lithornis and Promusophaga – paleognaths
  • Parvigyps – perhaps a diurnal raptor
  • Pediorallus – a paleognath or landfowl
  • Percolinus – perhaps another landfowl
  • "Precursor" – apparently a chimera of Charadriiformes and Psittaciformes (and possibly other) bones
  • Primapus – a swift-like bird
  • Primodroma – a tubenose, possibly a storm-petrel
  • Prophaeton – a tropicbirds relative
  • Proherodius – another waterbird
  • Proplegardis – a stork or ibis
  • Pulchrapollia – a parrot relative
  • Stintonornis – probably a hawk relative
  • Reptiles

    Crocodylians

  • Diplocynodon – an alligatoroid
  • Kentisuchus spenceri – a crocodylid
  • Snakes

  • Palaeophis toliapicus and P. typhaeus
  • Turtles and tortoises

  • Allaeochelys – a pig-nosed turtle
  • Argillochelys, Eochelone, Puppigerus and "Thalassochelys" sp. – true sea-turtles
  • Chrysemys bicarinata and C. testudiniformis – pond turtles
  • Eosphargis – a leatherback sea-turtle
  • Homopus comptoni – a tortoise
  • Lytoloma crassicostatum and L. planimentum – prehistoric sea-turtles
  • Palaeaspis – an African sideneck turtle
  • Podocnemis bowerbanki – an American sideneck turtle
  • Trionyx pustulatus and T. sp. – softshell turtles
  • Dacochelys and Pseudotrionyxincertae sedis
  • Bony fish
  • Acestrus elongatus, A. ornatus, Aglyptorhynchus sulcatus, A. venablasi, Xiphiorhynchus parvus and X priscus – swordfish relatives
  • Acipenser toliapicus – a true sturgeon
  • Albula oweni – a bonefish
  • Ampheristus toliapicus – a scorpionfish
  • Ardiodus marriottiincertae sedis
  • Argillichthys toombsi – a lizardfish relative
  • Aulopopsis depressifrons, A. egertoni and Labrophagus esocinus – flagfins
  • Beerichthys ingens and B. sp. – Two species of luvar or luvar-like fish
  • Bramoides brieni and Goniocranion arambourgi – pomfrets
  • Brychaetus muelleri – an arowana
  • Bucklandium diluvii – a naked catfish
  • Cylindracanthus rectus and Hemirhabdorhynchus elliotti – Blochiidae
  • Cymbium proosti, Eocoelopoma colei, E. curvatum, E. gigas, E. hopwoodi, Eothynnus salmoneus, Scombramphodon crassidens, S. sheppeyensis, Scombrinus macropomus, S. nuchalis, Sphyraenodus priscus, Tamesichthys decipiens, Wetherellus brevior, W. cristatus, W. longior and Woodwardella patellifrons – mackerel and tuna relatives
  • Diodon sp. – a porcupinefish
  • Egertonia isodonta and Phyllodus toliapicus – Phyllodontidae
  • Elops sp., Esocelops cavifrons, Megalops oblongus, M. priscus, Promegalops sheppeyensis and P. signeuxae – ladyfish
  • Enniskillenus radiatus – acanthomorph
  • Eutrichurides winkleri – a cutlassfish
  • Halecopsis insignis – Halecopsidae
  • Laparon alticeps and Whitephippus tamesis – spadefish
  • Lehmanamia sheppeyensis – a bowfin
  • Myripristis toliapicus, Naupygus bucklandi and Paraberyx bowerbanki – soldierfish
  • Percostoma angustum, Plesioserranus cf. wemmeliensi and Serranopsis londinensis – groupers
  • Podocephalus curryi, P. nitidus, Sciaenuropsis turneri and Sciaenurus bowerbanki – porgies
  • Progempylus edwardsi – a snake mackerel
  • Pseudosphaerodon antiquus and P. navicularis – wrasses?
  • Pycnodus bowerbanki and P. toliapicus – Pychnodontidae
  • Rhinocephalus planiceps and Trichurides sagittidens – hakes
  • Rhynchorhinus branchialis and R. major – Eccelidae
  • Tetratichthys antiquitatis – a jack mackerel
  • Cartilaginous fish
  • Abdounia beaugi, Carcharhinus sp. and Physogaleus secundus – requiem sharks
  • Aetobatis irregularis, Burnhamia daviesi, Myliobatis dixoni, M. latidens, M. raouxi and M. toliapicus – eagle rays
  • Anomotodon sheppeyensis – a goblin shark
  • Carcharias hopei, Jaekelotodus trigonalis, Odontaspis winkleri, Palaeohypotodus rutoti and Striatolamia macrota – sand sharks
  • Edaphodon bucklandi and Elasmodus hunteri – chimaeras
  • Dasyatis davisi and D. wochadunensis – stingrays
  • Galeorhinus lefevrei, G. minor, G. recticonus, G. ypresiensis, Mustelus whitei and Triakis wardi – hound sharks
  • Heterodontus vincenti, H. wardenensis and H. woodwardi – bullhead sharks
  • Hexanchus agassizi, H. collinsonae, H. hookeri, Notorhynchus serratissimus and Weltonia burnhamensis – cow sharks
  • Isisteus trituratus and Squalus minor – dogfish sharks
  • Isurolamna affinis, Isurus nova, I. praecursor, Lamna inflata, L. lerichei, Otodus obliquus and Xiphodolamia eocaena – white sharks
  • Megascyliorhinus cooperi, Scyliorhinus casieri, S. gilberti, S. pattersoni and S. woodwardi – catsharks
  • Pararhincodon sp? – an indeterminate shark
  • Raja sp.? – an indeterminate ray
  • Squatina prima – an angel shark
  • Crustaceans

  • Lobsters and shrimp
  • Archiocarabus bowerbanki
  • Callianassa sp.
  • Homarus morrisi 
  • Hoploparia gammaroides
  • Linuparus eocenicus & L. scyllariformis
  • Scyllarides tuberculatus
  • Scyllaridia koenigi
  • Thenops scyllariformis
  • Barnacles
  • Arcoscapellum quadratum
  • Scalpellum minutum and S. quadratum
  • Crabs
  • Campylostoma mutatiforme
  • Cyclocorystes pulchellus
  • Dromilites bucklandi & D. lamarki
  • Glyphthyreus wetherelli
  • Goniochela angulata
  • Harpactoxanthopsis cf. quadrilo
  • Mithracia libinioides
  • Oediosoma ambigua
  • Portunites incerta & P. stintoni
  • Xanthilites bowerbanki
  • Zanthopsis bispinosa, Z. dufori, Z. leachei, Z. nodosa and Z. unispinosa
  • Mantis shrimp
  • Squilla wetherelli
  • Molluscs

    Cephalopods
  • Aturia ziczac, Cimomia imperialis, Deltoidonautilus sowerbyi, Euciphoceras regale, Eutrephoceras urbanum, Hercoglossa cassiniana and Simplicioceras centrale – nautiluses
  • Belopterina levesquei, Belosepia blainvillei and B. sepioidea – cuttlefish
  • Bivalves
  • Abra splendens – Semelidae
  • Amygdalum depressum and Modiolus tubicola – Mytilidae
  • Anomiidae

  • Anomia scabrosa – a jingle shell
  • Enigmonia aenigmatica – a jingle shell
  • Arca nitens, A. tumescens and Glycymeris wrigleyi – ark clams
  • Arctica planata – Arcticidae
  • Astarte davisi, A. filigera and A. rugata
  • Astartidae

  • Calpitaria sulcataria – a venus clam
  • Corbula globosa – Corbulidae
  • Cuspidaria inflata and C. lamallosa – Cuspidariidae
  • Nuculana amygdaloides and N. prisca – Nuculanidae
  • Lentipecten corneus and Pecten sp. – scallops
  • Nemocardium nitens and N. semigranulatum – Cardiidae
  • Nucula consors – Nuculidae
  • Ostrea sp. – a true oysters
  • Pinna affinis – a pen shell
  • Pleurolectroma media and Pteria papyracea – pearl oysters
  • Pycnodonte gryphovicina – Pycnodontidae
  • Teredina personata and Teredo sp. – shipworms
  • Thyasira angulata – Thyasiridae
  • Thyasira goodhali – Thyasiridae
  • Venericardia trinobantium – Carditidae
  • Verticordia sulcata – Verticordiidae
  • Gastropods
  • Acrilla cymaea, Foratiscala perforata, Litoriniscala scalaroides and Undiscala primaeva – wentletraps
  • Aporrhais sowerbii and Eotibia lucida – true conchs
  • Bathytoma granata, B. turbida, Clavatula conica, Cochlespira gyrata, Conolithus concinnus, Endiatoma cerithiformis, Fusiturris selysi, F. simillima, F. wetherelli, Gemmula koninckii, Hemipleurotoma fasciolata, H. prestwichi, Pseudotoma topleyi, Surculites errans, S. velatus, Turricula crassa, T. helix, T. latimarginata, T. nanodis, T. symmetrica and T. teretrium – Conoidea
  • Bonellitia clathratum and B. laeviuscula – nutmeg shells
  • Bullinella sp., Crenilabium elongatum, ?Roxiana sp., Scaphander ?parisiensis and Tornatellaea simulata – opisthobranchs
  • Camptoceratops prisca, Spiratella mercinensis, S. taylori and S. tutelina – sea-butterflies
  • Cassis striata and Mambrina gallica – tun shells
  • Cepatia cepacea, Daphnobela juncea, Litiopa sulculosa, Orthochetus elongatus and Stellaxis pulcherincertae sedis
  • Eocypraea oviformis – a cowrie
  • Euspira glaucinoides and Sinum clathratum – moon snails
  • Falsifusus londini, Fusinus coniferus, F. wetherelli, Pseudoneptunea curta, Siphonalia highgatensis, Streptolathyrus triliniatus, S. zonulatus, Wrigleya complanata and W. transversaria – true whelks
  • Ficopsis multiformis – a fig shell
  • Lacunella sp. – a periwinkle
  • Mathilda sororcula – Mathildidae
  • Murex subcristatus and Paziella argillacea – murex snails
  • Pachysyrnola sp. and Turbonilla subterranea – pyramid shells
  • Patella sp. – Patellidae
  • Ptychatractus aff. interuptus, Scaphella wetherelli and Volutospina nodosa – volutes
  • Rilla cf. tenuistriata – Streptaxidae
  • Ringicula turgida – Ringiculidae
  • Sassia morrisi – a triton shell
  • Sigapatella sp. – Calyptraeidae
  • Tornus sp. and Turboella cf. misera – Rissoidae
  • Xenophora extensum – a carrier shell
  • Tusk shells
  • Antalis anceps and A. nitens
  • Echinoderms

  • Asteropecten crispatus, Coulonia colei, Hemiaster bowerbanki, Hippasteria tuberculata, Ophioglypha wetherelli and Teichaster stokesii – starfish
  • Coelopleurus wetherelli, Micraster sp. and Schizaster sp. – sea urchins
  • Democrinus londinensis – crinoid
  •  ?Ophiacantha sp., Ophioglypha wetherelli, Ophiomusium sp. and Ophiura wetherelli – brittlestars
  • Annelids

  • Rotularia bognoriensis
  • Serpula trilineata
  • Cnidarians

  • Paracyathus brevis and P. caryophyllus – corals
  • Graphularia wetherelli – hydrozoan
  • Other invertebrates

  • Adenellopsis wetherelli, Aimulosia sp., Batopora clithridiata, Beisselina sp., Cribrilina sp., Didymosella sp., Dittosaria wetherelli, Entalophora sp., Idmonia sp., Lunulites sp., Nellia sp., Pachythecella incisa, Vibracellina sp. and Websteria crissioides – bryozoans
  • Hemiptera gen. et sp. indet. – true bug
  • Lingula tenuis, Terebratulina striatula and T. wardenensis – lampshells
  • Stelleta sp. – sponge
  • Ichnofossils

  • Ditrupa plana – polychaete worm tubes?
  • Scolithos
  • References

    London Clay Wikipedia