Protungulatum

Protungulatum ('first ungulate') is an extinct genus of mammal first found in the Bug Creek Anthills in northeastern Montana. The Bug Creek Anthills were initially believed to be Late Cretaceous (latest Maastrichtian) because of the presence of the remains of non-avian dinosaurs and common Cretaceous mammals, but these were later shown to have been reworked from Late Cretaceous strata, and consequently the Bug Creek Anthills are currently believed to be Early Paleocene (Puercan) in age. Remains from the Ravenscrag Formation of Saskatchewan, Canada have been assigned to P. donnae. These remains may also be Cretaceous in age, but the age of the Ravenscrag Formation is not entirely certain. In 2011, remains of a new species of Protungulatum, P. coombsi, from the Hell Creek Formation, which is definitely Late Cretaceous (Maastrichtian) in age, proved that Protungulatum was present in both the Cretaceous and the Paleocene. It was initially assigned to the order condylarthra, a group of archaic "ungulates", that is now known to be polyphyletic. According to Archibald et al. (2011), Protungulatum is not even definitely a placental mammal. Some studies have found it to be close to Cetartiodactyla, but the most recent analysis helds it to be a non-placental eutherian.

Contents

• Protungulatum top 6 facts
• Phylogeny
• References

The date of divergence of Protungulatum and Placentalia is approximately 75 Ma.