Girish Mahajan (Editor)

Piperaceae

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Kingdom
  
Clade
  
Scientific name
  
Piperaceae

Rank
  
Family

Clade
  
Order
  
Higher classification
  
Piperaceae Piperaceae species list Pepper of Hawaii Tropical Botanical Garden

Lower classifications
  
Black pepper, Radiator plants, Long pepper, Kava, Pepper

Peperomia argyreia piperaceae watermelon peperomia


The Piperaceae, also known as the pepper family, are a large family of flowering plants. The group contains roughly 3,600 currently accepted species in 13 genera. The vast majority of peppers can be found within the two main genera: Piper (2000 species) and Peperomia (1600 species).

Contents

Piperaceae Flowering Plant Families UH Botany

Members of the Piperaceae may be small trees, shrubs, or herbs. The distribution of this group is best described as pantropical.

Piperaceae Macropiper excelsum Piperaceae image 21720 at PhytoImagessiuedu

The best-known species is Piper nigrum, which yields most peppercorns that are used as spices, including black pepper, although its relatives in the family include many other spices.

Piperaceae Angiosperm families Piperaceae CA Agardh

Pepper vines from piperaceae family grow in kerala india


Etymology

Piperaceae wwwbotanyhawaiiedufacultycarrimagespeptet

The name Piperaceae is likely derived from the Sanskrit term pippali, which was used to describe long peppers (like those of Piper longum).

Taxonomy

The APG III system of 2009 recognizes this family, and assigns it to the order Piperales in the unranked clade magnoliids. The family consists of five genera: Piper, Peperomia, Zippelia, Manekia, and Verhuellia. The previously recognised Pacific genus Macropiper, was recently merged into Piper. A tentative cladogram showing relationships based on Wanke et al. (2007) is shown below. This phylogeny was based on 6000 basepairs of chloroplast DNA. Only recently has it become clear that Verhuellia is sister to the other four genera in the family.

Characteristics

Members of pepper family are small trees, shrubs, or perennial or annual herbs.

Roots and stems

Plants are often rhizomatous, and can be terrestrial or epiphytic. The stems can be either simple or branched.

Leaves

Piperaceae PIPERACEAE Pybio Paraguay Biodiversidad

Leaves are simple with entire margins, and are positioned at the base of the plant or along the stem, and can be alternate, opposite, or whorled in arrangement. Stipules are usually present, as are petioles. The leaves are often noticeably aromatic when crushed.

Flowers

Inflorescences (in the form of spikes) are terminal, opposite the leaves, or located in the axils. Flowers are bisexual, with no perianth, each flower is subtended by a peltate bract. Stamens are 2-6, and hypogynous, with 2-locular anthers. There are usually 3-4 stigmas attached to a single pistil per flower, which is 1 or 3-4 carpellate. The ovary is 1 locular, and superior.

Fruits and seeds

Fruits are drupelike, with a single seed per fruit. The seeds have a minute embryo, and mealy perisperm.

Genera

Subfamily Verhuellioideae Samain & Wanke

  • Verhuellia Miquel 1843 (three species)
  • Subfamily Zippelioideae Samain & Wanke

  • Zippelia Blume 1830 (one species)
  • Manekia Trelease 1927 (six species)
  • Subfamily Piperoideae Arnott

  • Piper Linnaeus 1753 (about 2000 species)
  • Peperomia Ruiz & Pavon 1794 (about 1600 species)
  • References

    Piperaceae Wikipedia