Neuquensaurus

HistoryEdit

In 1893, Richard Lydekker named Titanosaurus australis, based on a series of caudal vertebrae and limb elements. The remains had been found by Santiago Roth and F. Romero in the Neuquén Province of Argentina at the Neuquén River, and were by Lydekker assigned to a single individual. Six caudal vertebrae, with the inventory numbers MLP Ly 1-6-V-28-1, were the holotype of the species. They had probably been found in a layer of the Anacleto Formation.

Some elements that had been referred to Titanosaurus australis were reassigned to Laplatasaurus araukanicus by Friedrich von Huene in 1929. The same year, von Huene named a Titanosaurus robustus, and claimed it differed from T. australis in the limb material. Von Huene described all the slender limb material to T. australis, but did not identify any differentiating features between the vertebrae. When describing T. robustus, von Huene did not really compare other genera and species to it. From the syntype material assigned by von Huene to T. robustus José Fernando Bonaparte et al. in 1978 chose four lectotypes, specimens MLP 26-250, MLP 26-252, MLP 26-254, and MLP 26-259, a left femur, both ulnae, and a left radius.

In 1986, Jaime Eduardo Powell, concluding that Titanosaurus australis was less similar than Laplatasaurus araukaicus to Titanosaurus indicus, named a separate genus: Neuquensaurus. However, he did so in an unpublished dissertation which caused Neuquensaurus australis and Neuquensaurus rubustus to remain invalid nomina ex dissertatione. In 1990, the two species were assigned to Saltasaurus by John Stanton McIntosh, as a Saltasaurus australis and a Saltasaurus robustus, claiming that the features found by Bonaparte were not of sufficient taxonomic importance to justify a generic separation.

In 1992 Powell validly named Neuquensaurus, with as type species Titanosaurus australis of which the combinatio nova then is Neuquensaurus australis. He also found Titanosaurus robustus to be assignable to the new genus, but considered it non-diagnostical, and so a nomen dubium.

DescriptionEdit

This dinosaur is believed to have possessed armor-like osteoderms. A relatively small sauropod, with a femur only 0.75 metres (2.5 ft) long. It is one of the most completely known of Patagonian sauropods. In addition to the original fossils described by Lydekker in 1893, it is represented by fossils collected in the early twentieth century, and more recent material, including a well preserved and partially articulated specimen described in 2005 (with two associated osteoderms), as well as additional undescribed material.

SkeletonEdit

Neuquensaurus has an almost complete skeleton. Among the preserved elements are a scapula. The scapula is co-ossified with the coracoid, a feature also found in Opisthocoelicaudia. In general aspect, the scapula resembles Saltasaurus, Opisthocoelicaudia, Lirainosaurus, and Alamosaurus. The coracoids of the scapulae are roughly quadrangular in shape, and are alike Saltasaurus and Lirainosaurus, but not Opisthocoelicaudia, Rapetosaurus, or Isisaurus, all of which have a rounded coracoid.

Distinguishing characteristicsEdit

Many characters distinguish Neuquensaurus from other titanosaurians. The features found by Otero in 2010 are listed below:

ClassificationEdit

Neuquensaurus is a derived saltasaurine. Its closest relatives were Saltasaurus, Rocasaurus, and Bonatitan, and together they made up Saltasaurinae.

PaleoecologyEdit

Neuquensaurus is known from fossils from the Anacleto Formation.