The Nemastomatidae are a family of harvestmen with about 170 described species in 16 recent genera. Several fossil species and genera are known.
Unlike some related currently recognized families, the Nemastomatidae are monophyletic.
Members of the Nemastomatidae range in body length from about one to almost six millimeters. Their chelicerae are of normal proportians, but the pedipalps are very elongated and thin in some groups. Leg length is likewise variable.
The subfamily Ortholasmatinae (ten species in two genera) occur on both sides of the Pacific Ocean: in western North America from Mexico to British Columbia and eastern Asia (Japan and northern Thailand. The other subfamily, Nemastomatinae, occur all over Europe up to Iceland and the Caucasus, in the Atlas Mountains of North Africa, from Anatolia to northern Iran, with a few species found outside this region, in Central Asia and the Himalayas. Most species are restricted to very small mountainous southern regions.
The Nemastomatidae are probably a sister group to the genera Dicranolasma (Dicranolasmatidae) and Trogulus (Trogulidae).
The genus name Nemastoma is a combination of Ancient Greek nema "thread" and stoma "mouth", referring to the elongated pedipalps.
Buresiolla Kratochvíl & Miller, 1958
Buresiolla bureschi (Roewer, 1926)
Buresiolla abchasica Ljovuschkin & Starobogatov, 1963 — Abkhazia
Buresiolla caecum (Nowikoff, 1931) — Crimea
Nemaspela Silhavý, 1966
Nemaspela sokolovi (Ljovuschkin & Starobogatov, 1963)
Nemaspela birsteini Levushkii, 1972
Nemaspela taurica (Lebedinskiy, 1914) — Crimea
Giljarovia Kratochvíl & Miller, 1958
Giljarovia rossica Kratochvíl & Miller, 1958
Giljarovia stridula (Kratochvíl & Miller, 1958)
Giljarovia turcica Gruber, 1976
Nemastoma C. L. Koch, in Hahn & C. L. Koch 1836
subgenus Nemastoma
Nemastoma aeginum Roewer, 1951
Nemastoma amulleri Roewer, 1951
Nemastoma anatolicum Roewer, 1962
Nemastoma armeniacum Roewer, 1951
Nemastoma atticum Roewer, 1927
Nemastoma bacilliferum Simon, 1879 — Spain
Nemastoma bacilliferum bacilliferum Simon, 1879 — Pyrenees
Nemastoma bacilliferum simoni Roewer, 1914
Nemastoma bacilliferum simplex Simon, 1913
Nemastoma bacilliferum dipentatum Rambla, 1959
Nemastoma bidentatum Roewer, 1914 — central Europe
Nemastoma bidentatum bidentatum Roewer, 1914
Nemastoma bidentatum sparsum Gruber & Martens, 1968
Nemastoma bidentatum relictum Gruber & Martens, 1968
Nemastoma bimaculosum Roewer, 1951
Nemastoma brevipalpatum Roewer, 1951
Nemastoma caecum Grese, 1911 — Crimea
Nemastoma cancellatum Roewer, 1917 — Bosnia
Nemastoma caporiaccoi Roewer, 1951
Nemastoma carbonarium Simon, 1907 — Spain
Nemastoma carpathicum Roewer, 1951
† Nemastoma clavigerum Menge, 1854 — fossil
Nemastoma corcyraeum Roewer, 1917 — Korfu
Nemastoma cypricum Roewer, 1951
† Nemastoma denticulatum Koch & Berendt, 1854 — Baltic amber fossil: Oligocene
Nemastoma dentigerum Canestrini, 1873 — Italy
Nemastoma dubium Mello-Leitão, 1936 — Spain
Nemastoma emigratum Roewer, 1959
Nemastoma ferkeri Roewer, 1951
Nemastoma filipes Roewer, 1917 — Transcaucasus
Nemastoma formosum Roewer, 1951
Nemastoma franzi Kraus, 1959
Nemastoma funebre Redikorzev, 1936 — Russia
Nemastoma gallwitzi Roewer, 1923 — Serbia
Nemastoma gigas Sørensen, in Lendl 1894 — Hungary
Nemastoma globuliferum L. Koch, 1867
Nemastoma gostivarense Hadzi, 1973
Nemastoma gracile Redikorzev, 1936 — Russia
Nemastoma hankiewiczii Kulczynski, 1909 — Portugal
Nemastoma hermanni Kraus, 1959
Nemastoma ikarium Roewer, 1951
Nemastoma insulare Roewer, 1951
Nemastoma ios Roewer, 1917 — Sporaden
Nemastoma kastneri Roewer, 1951
Nemastoma lessinensis Caporiacco, 1940 — Italy
Nemastoma leiobunum Wankel, 1851
Nemastoma lilliputanum (Lucas, 1847) — Algeria
Nemastoma lindbergi Roewer, 1959
Nemastoma longipalpatum Roewer, 1951
Nemastoma longipes Schenkel, 1947 — Albania
Nemastoma maarebense Simon, 1913 — Algeria
Nemastoma machadoi Roewer, 1951
Nemastoma mackenseni Roewer, 1923 — Serbia
Nemastoma macedonicum Hadzi, 1973
Nemastoma manicatum Simon, 1913 — Spain
Nemastoma moesiacum Roewer, 1917 — Serbia and Bosnia
Nemastoma monchiquense Kraus, 1961
Nemastoma montenegrinum Nosek, 1904 — Montenegro
Nemastoma navarrense Roewer, 1951
Nemastoma nervosum Roewer, 1923 — Serbia
Nemastoma perfugium Roewer, 1951
Nemastoma polonicum Roewer, 1951
Nemastoma pyrenaeum Simon, 1879 — Pyrenees
Nemastoma reimoseri Roewer, 1951
Nemastoma rhinoceros Roewer, 1917 — Spain
Nemastoma riparium Roewer, 1951
Nemastoma romanium Roewer, 1951
Nemastoma rude Simon, 1881 — Alps
Nemastoma santorinum Roewer, 1951
Nemastoma scabriculum Simon, 1879 — Pyrenees
Nemastoma schenkeli Roewer, 1951
Nemastoma schuelleri Gruber & Martens, 1968
Nemastoma senussium Roewer, 1951
Nemastoma sexmucronatum Simon, 1911 — Spain
Nemastoma simplex Giltay, 1933 — Balkans
Nemastoma spinosissima Kraus, 1961
Nemastoma spinulosum L. Koch, 1869 — Greece
Nemastoma strasseri Roewer, 1951
Nemastoma stussineri Simon, 1885 — Greece
† Nemastoma succineum Roewer, 1939 — fossil: Baltic amber
Nemastoma tenebricosum Redikorzev, 1936 — Russia
Nemastoma tenue Hadzi, 1973
Nemastoma titaniacum Roewer, 1914 — Bosnia
Nemastoma transsylvanicum Gruber & Martens, 1968
Nemastoma triste (C. L. Koch, 1835) — central Europe
Nemastoma triste triste C. L. Koch, in Herrich-Schaeffer 1835
Nemastoma troglodytes (Wankel, 1861)
† Nemastoma tuberculatum Koch & Berendt, 1854 — fossil: Oligocene
Nemastoma tunetanum Roewer, 1951
Nemastoma vitynae Roewer, 1927
subgenus Dromedostoma
Nemastoma acrospinosum Roewer, 1951
Nemastoma acrospinosum acrospinosum Roewer, 1951
Nemastoma acrospinosum pretneri Hadzi, 1973
Nemastoma bipunctatum Hadzi, 1973
Nemastoma bolei Hadzi, 1973
Nemastoma corneluttii Hadzi, 1973
Nemastoma grabovicae Hadzi, 1973
Nemastoma megarae Hadzi, 1973
Nemastoma multisignaltum Hadzi, 1973
Nemastoma nigrum Hadzi, 1973
Nemastoma radewi Roewer, 1926 — cave in Balkans
Nemastoma ryla Roewer, 1951
Nemastoma sketi Hadzi, 1973
subgenus Lugubrostoma
Nemastoma boenicum
Nemastoma goliae Hadzi, 1973
Nemastoma lugubre (Müller, 1776)
Nemastoma lugubre lugubre (Müller, 1776)
Nemastoma lugubre bimaculatum (Fabricius, 1775) — Europe
Nemastoma lugubre unicolor Roewer, 1914 — southern Europe
Nemastoma sarae Hadzi, 1973
Paranemastoma Redikorzew, 1936
Paranemastoma bicuspidatum (C. L. Koch, 1835)
Paranemastoma ancae Avram, 1973
Paranemastoma charitonovi (Mcheidze, 1952)
Paranemastoma kalischevskyi (Roewer, 1951)
Paranemastoma kochii (Nowicki, 1870)
Paranemastoma quadripunctatum (Perty, 1833) — Europe
Paranemastoma quadripunctatum quadripunctatum (Perty, 1833)
Paranemastoma quadripunctatum aurosum L. Koch, 1869
Paranemastoma quadripunctatum armatum Kulczynski, 1909 — southeastern Europe
Paranemastoma quadripunctatum thessalum Simon, 1885 — Greece
Paranemastoma quadripunctatum werneri Kulczynski, 1903 — Asia Minor
Paranemastoma quadripunctatum humerale C. L. Koch, 1839 — southeastern Europe
Paranemastoma roeweri Starega, 1978
Paranemastoma sillii (Hermann, 1871) — central Europe
Paranemastoma suchumium (Roewer, 1951)
Paranemastoma supersum (Roewer, 1951)
Paranemastoma umbo (Roewer, 1951)
Paranemastoma aurigerum (Roewer, 1951)
Paranemastoma aurigerum aurigerum Roewer, 1951
Paranemastoma aurigerum joannae Starega, 1976
Histricostoma Kratochvíl & Miller, 1958
Histricostoma drenskii Kratochvíl & Miller, 1958
Histricostoma dentipalpe (Ausserer, 1867) — central Europe
Histricostoma argenteolunulatum (Canestrini, 1872) — Corcica, Italy
Histricostoma caucasicum (Redikorzev, 1936) — Russia
Carinostoma Kratochvíl & Miller, 1958
Carinostoma carinatum (Roewer, 1914) — southeastern Europe
Carinostoma elegans (Sørensen, 1894) — Hungary
Carinostoma elegans elegans (Sørensen, 1894)
Carinostoma elegans batorligetiense (Szalay, 1951)
Carinostoma ornatum (Hadzi, 1940)
Centetostoma Kratochvíl & Miller, 1958
subgenus Centetostoma
Centetostoma centetes (Simon, 1881) — Alps
subgenus Cretostoma
Centetostoma creticum (Roewer, 1927)
Mitostoma Roewer, 1951
Mitostoma armatissimum Roewer, 1962
Mitostoma daccordii Tedeschi & Sciaky, 1997
Mitostoma fabianae Tedeschi & Sciaky, 1997
Mitostoma sabbadinii Tedeschi & Sciaky, 1997
Mitostoma orghidani Avram, 1969
Mitostoma dumitrescui Avram, 1969
Mitostoma patrizii Roewer, 1958
Mitostoma saxonicum
Mitostoma asturicum Roewer, 1951
Mitostoma omalosum Roewer, 1951
Mitostoma chrysomelas (Hermann, 1804) — Europe
Mitostoma chrysomelas chrysomelas (Hermann, 1804)
Mitostoma chrysomelas michielii Hadzi, 1973
Mitostoma chrysomelas multidenticulatum Hadzi, 1973
Mitostoma chrysomelas alpinum Hadzi, 1931 — Yugoslavia
Mitostoma anophthalmum (Fage, 1946) — Italy
Mitostoma carneluttii Hadzi, 1973
Mitostoma helenae Avram, 1970
Mitostoma macedonicum Hadzi, 1973
Mitostoma moldavica Avram, 1970
Mitostoma rodnae Avram, 1970
Mitostoma zmajevicae Hadzi, 1973
Mitostoma olgae Silhavý, 1946 — Dalmatia, Montenegro
Mitostoma olgae olgae Silhavý, 1946
Mitostoma olgae decorum Silhavý, 1946
Mitostoma olgae kratochvili Silhavý, 1946
Mitostoma olgae zorae Hadzi, 1973
Mitostoma valdemonense Marcellino, 1977
Dendrolasma Banks, 1894
Dendrolasma mirabile Banks, 1894 — Washington
Dendrolasma angka P. J. Schwendinger & J. Gruber, 1992 — Thailand
Dendrolasma dentipalpe W. A. Shear & J. Gruber, 1983 — California
Dendrolasma parvula (Suzuki, 1963)
† Halitherses Giribet & Dunlop, 2005
† Halitherses grimaldii Giribet & Dunlop, 2005 — fossil: Cretaceous (Albian) Burmese amber, c. 100 mya
Ortholasma Banks, 1894
Ortholasma bolivari (Goodnight & Goodnight, 1942) — Mexico
Ortholasma coronadense Cockerell, 1916 — Coronados Islands
Ortholasma levipes W. A. Shear & J. Gruber, 1983 — California
Ortholasma pictipes Banks, 1911 — California
Ortholasma rugosum Banks, 1894 — California
Ortholasma sbordonii Silhavý, 1973
Nemastomella Mello-Leitão, 1936
Nemastomella integripes Mello-Leitão, 1936 — Spain
Hadzinia Silhavý, 1966
Hadzinia karamani (Hadzi, 1940)
Acromitostoma Roewer, 1951
Acromitostoma hispanum (Roewer, 1917) — Spain: Granada
Acromitostoma rhinoceros Roewer, 1951
Mediostoma Kratochvíl & Miller, 1958
Mediostoma graecum (Roewer, 1917)
Mediostoma topolium (Roewer, 1951)
Mediostoma pamiricum W. Starega, 1986 — Tajikistan
Mediostoma ceratocephalum Gruber, 1976
Pyza Starega, 1976
Pyza bosnica (Roewer, 1916)
† Rhabdotarachnoides Haupt, 1956 — fossil
† Rhabdotarachnoides simoni Haupt, 1956 — Permian