Kingdom Animalia Phylum Arthropoda Order Nektaspida | Stem group Chelicerata Rank Family | |
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Similar Misszhouia, Naraoia, Buenaspis, Tegopelte, Leanchoilia | ||
Naraoiidae how to pronounce it
Naraoiidae is a family, of extinct, soft-shelled trilobite-like arthropods, that belongs to the order Nectaspida. Species included in the Naraoiidae are known from the second half of the Lower Cambrian to the end of the Upper Silurian. The total number of collection sites is limited and distributed over a vast period of time: Maotianshan Shale and Balang Formation (China), Burgess Shale and Bertie Formation (Canada), the Šárka Formation (Czech Republic), Emu Bay Shale (Australia), Idaho and Utah (USA). This is probably due to the rare occurrence of the right circumstances for soft tissue preservation, needed for these non-calcified exoskeletons.
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Ecology
Naraoiids probably were deposit feeders (Naraoia and Pseudonaraoia), predators or scavengers (Misszhouia), living on the sea floor.
Description
The species of the family Naraoiidae are almost flat (dorso-ventrally). The upper (or dorsal) side of the body consists of a non-calcified transversely oval or semi-circular headshield (cephalon), and a circular to long oval tailshield (pygidium) equal to or longer than the cephalon, without any body segments in between. The body is narrowed at the articulation between cephalon and pygidium. The antennas are long and many-segmented. There are no eyes. The 17 to 25 pairs of legs have two branches on a common basis, like trilobites. The outer (dorsal) branches of the limbs (exopods) have flattened side branches (setae) on the shaft (probably acting as gills). The inner branches (or endopods are composed of 6 or 7 segments (or podomeres).
Differences with other Nectaspida
Naraoiidae lack thoracic segments (or tergites), while the species of the sister family Liwiidae have between 3 and 6 tergites.
Taxonomic history
The taxonomic placement of the Naraoiidae has long been debated until detailed appendages were uncovered, that showed that N. compacta shares biramous legs of very comparable anatomy with trilobites. Some debate is still going on if the parent taxon Nektaspida should be included in the Trilobita, or is better placed as a sister group.