Growing degree days (GDD), also called growing degree units (GDUs), are a heuristic tool in phenology. GDD are a measure of heat accumulation used by horticulturists, gardeners, and farmers to predict plant and animal development rates such as the date that a flower will bloom, an insect will emerge from dormancy, or a crop will reach maturity.
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Introduction
In the absence of extreme conditions such as unseasonal drought or disease, plants grow in a cumulative stepwise manner which is strongly influenced by the ambient temperature. Growing degree days take aspects of local weather into account and allow gardeners to predict (or, in greenhouses, even to control) the plants’ pace toward maturity.
Unless stressed by other environmental factors like moisture, the development rate from emergence to maturity for many plants depends upon the daily air temperature. Because many developmental events of plants and insects depend on the accumulation of specific quantities of heat, it is possible to predict when these events should occur during a growing season regardless of differences in temperatures from year to year. Growing degrees (GDs) is defined as the number of temperature degrees above a certain threshold base temperature, which varies among crop species. The base temperature is that temperature below which plant growth is zero. GDs are calculated each day as maximum temperature plus the minimum temperature divided by 2 (or the mean temperature), minus the base temperature. GDUs are accumulated by adding each day’s GDs contribution as the season progresses.
GDUs can be used to: assess the suitability of a region for production of a particular crop; estimate the growth-stages of crops, weeds or even life stages of insects; predict maturity and cutting dates of forage crops; predict best timing of fertilizer or pesticide application; estimate the heat stress on crops; plan spacing of planting dates to produce separate harvest dates. Crop specific indices that employ separate equations for the influence of the daily minimum (nighttime) and the maximum (daytime) temperatures on growth are called crop heat units (CHUs).
GDD calculation
GDD are calculated by taking the integral of warmth above a base temperature, Tbase (usually 10 °C):
A simpler, approximately equivalent formulation uses the average of the daily maximum and minimum temperatures compared to a Tbase. As an equation:
If the mean daily temperature is lower than the base temperature then GDD=0.
GDDs are typically measured from the winter low. Any temperature below Tbase is set to Tbase before calculating the average. Likewise, the maximum temperature is usually capped at 30 °C because most plants and insects do not grow any faster above that temperature. However, some warm temperate and tropical plants do have significant requirements for days above 30 °C to mature fruit or seeds.
For example, a day with a high of 23 °C and a low of 12 °C (and a base of 10 °C) would contribute 7.5 GDDs.
A day with a high of 13 °C and a low of 10 °C (and a base of 10 °C) would contribute 1.5 GDDs.
Pest control
Insect development and growing degree days are also used by some farmers and horticulturalists to time their use of organic or biological pest control or other pest control methods so they are applying the procedure or treatment at the point that the pest is most vulnerable. For example:
Honeybees
Several beekeepers are now researching the correlation between growing degree days-GDD and the lifecycle of a honeybee colony.{http://www.beeculture.com/phenology-and-its-value-to-beekeepers/} {http://bee-quick.com/reprints/budding.pdf} {https://u.osu.edu/beelab/phenology-for-beekeepers/}
Baselines
10 °C is the most common base for GDD calculations, however, the optimal base is often determined experimentally based on the lifecycle of the plant or insect in question.
GDDs may be calculated using either Celsius or Fahrenheit, though they must be converted appropriately; for every 9 GDDF there is 5 GDDC, or in conversion calculation:
GDDC = 5/9 * GDDF