Rahul Sharma (Editor)

Elymniini

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Kingdom
  
Animalia

Order
  
Lepidoptera

Family
  
Nymphalidae

Scientific name
  
Elymniini

Rank
  
Tribe

Phylum
  
Arthropoda

Superfamily
  
Papilionoidea

Subfamily
  
Satyrinae

Higher classification
  
Satyrinae

Elymniini httpsd1k5w7mbrh6vq5cloudfrontnetimagescache

Similar
  
Nymphalidae, Satyrinae, Butterflies and moths, Butterfly, Kirinia

The Elymniini (true browns) are one of the large tribes of the browns subfamily (Satyrinae) in the brush-footed butterfly family (Nymphalidae). Sometimes, they are elevated to subfamily status as Elymniinae.

Contents

Systematics

They are commonly subdivided into four subtribes (tribes when considered a subfamily). The largest tribe is called Lethina, with Parargina usually treated as a junior synonym. But this is not correct as it seems: according to cladistic analysis of mtDNA COI and NADH dehydrogenase subunit 1 sequence data, the Parargina – i.e. the group around Pararge, including at least the very similar Lasiommata as well as Tatinga and it is the doubtfully distinct relative Chonala – form a distinct lineage after all, immediately basal to the distinctly apomorphic Mycalesina and the more plesiomorphic Lethina proper; the accurate circumscription of these subtribes remains yet undetermined. In any case, these three constitute the bulk of the tribe's diversity. The Elymniina and Zetherina, by contrast, quite certainly form a less diverse clade that diverged early on in the evolution of the tribe.

Selected genera

Subtribes are listed in the presumed phylogenetic sequence, from the most ancient to the most advanced; genera are listed alphabetically. Some species are also listed.

Subtribe Elymniina

  • Elymnias Hübner, 1818 – palmflies
  • Elymnias hypermnestra – Common Palmfly
  • Elymniopsis Fruhstorfer, 1907
  • Hyantis Hewitson, 1862
  • Morphopsis Oberthür, 1880
  • Subtribe Zetherina

  • Callarge Leech, 1892
  • Ethope Moore, 1866 (sometimes placed in Lethina)
  • Neorina Westwood, 1850 (sometimes erroneously placed in Lethina)
  • Neorina patria – White Owl
  • Penthema Doubleday, 1848
  • Zethera Felder, 1861
  • Subtribe Lethina (including Parargina, but paraphyletic)

  • Chonala Moore, 1893 (might belong in Tatinga)
  • Enodia Hübner, [1819] – pearly-eyes
  • Enodia anthedon – Northern Pearly-eye
  • Hanipha Moore, 1880
  • Kirinia Moore, 1893 (including Esperarge)
  • Lasiommata Westwood, 1841 – walls and wallbrowns (formerly in Pararge)
  • Lasiommata maera – Large Wall, Large Wall Brown
  • Lasiommata megera – Wall Brown, Wall
  • Lasiommata menava – Dark Wall
  • Lasiommata petropolitana – Northern Wall Brown
  • Lasiommata schakra – Common Wall
  • Lethe – treebrowns, woodbrowns, foresters and relatives
  • Lopinga Moore, 1893 (formerly in Pararge)
  • Lopinga achine – Woodland Brown
  • Mandarinia Leech, 1892
  • Neope Moore, 1866
  • Ninguta Moore, 1892
  • Nosea Koiwaya, 1993
  • Orinoma Gray, 1846
  • Pararge Hübner, 1819
  • Pararge aegeria – Speckled Wood
  • Ptychandra C. & R. Felder, 1861
  • Rhaphicera Butler, 1867
  • Rhaphicera moorei – Small Tawny Wall
  • Satyrodes Scudder, 1875
  • Satyrodes eurydice – Eyed Brown
  • Tatinga Moore, 1893 (might include Chonala)
  • Subtribe Mycalesina

  • Bicyclus Kirby, 1871
  • Bletogona C. & R. Felder, 1867
  • Hallelesis Condamin, 1961
  • Heteropsis Westwood, 1850 (including Admiratio, Henotesia, Houlbertia, Masoura)
  • Mycalesis – bushbrowns
  • Nirvanopsis
  • Orsotriaena – Smooth-eyed Bush-brown
  • Pseudomycalesis Tsukada & Nishiyama, 1979
  • Fossil record

    Several prehistoric genera have been described from fossils:

  • Neorinella Martins Neto et al., 1993
  • Neorinopsis
  • Pseudoneorina Nel & Descimon, 1986
  • Satyrites Scudder, 1872
  • They are usually all assumed to belong to the Lethina; due to the non-monophyly of that group, all that can be said is that they are probably fairly basal members of the more diverse radiation. Fossils from near the time of such divergences often yield interesting insight into the patterns of evolution.

    References

    Elymniini Wikipedia


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