Arachniodes species include but are not limited to those in the following list. A species name preceded by (=) is considered by the referenced authority to be a synonym of the accepted species name listed above it.
Carl Ludwig von Blume published a description of the first Arachniodes species in Enumeratio Plantarum Javae et Insularum Adjacentium (An Enumeration of the Plants of Java and Adjacent Islands) in 1828.
Arachniodes aspidioides Blume 1828 The next Arachniodes species descriptions were published by Mary Douglas Tindale in Contributions from the New South Wales National Herbarium , volume 3, number 1, in 1961. This transferred two species from Conrad Vernon Morton's one-year-old Byrsopteris genus to Blume's 133-year-old Arachniodes genus.
Arachniodes amabilis (Blume) Tindale 1961 (India, Japan, Nepal, Philippines, Sri Lanka, Guangdong, Sichuan, Taiwan, Yunnan)Arachniodes aristata (G.Forst.) Tindale 1961 – East Indian holly fern (India, Japan, South Korea, Sri Lanka)Jisaburo Ohwi published the following Arachniodes species in the Journal of Japanese Botany , volume 37, number 3, in 1962.
Arachniodes assamica (Kuhn) Ohwi 1962 (Tibet, Yunnan, India, Nepal)Arachniodes cavalerii (H.Christ) Ohwi 1962 Arachniodes maximowiczii (Baker) Ohwi 1962 Arachniodes miqueliana (Maxim. ex Franch. & Sav.) Ohwi 1962 Arachniodes mutica (Franch. & Sav.) Ohwi 1962 Arachniodes nipponica (Rosenst.) Ohwi 1962 Arachniodes pseudoaristata (Tagawa) Ohwi 1962 Arachniodes simplicior (Makino) Ohwi 1962 – simpler holly fernArachniodes sinomiqueliana (Ching) Ohwi 1962 Arachniodes standishii (T.Moore) Ohwi 1962 – upside-down fernThe following species were transferred into Arachniodes by Ren Chang Ching in an article entitled "On the Nomenclature of the Compound-leaved Polysticha", published in the journal Acta Botanica Sinica , volume 10, number 3, Peking, China, September, 1962. Besides those listed below, a number of his species had already been published by Ohwi a few months earlier, and so appear in the section above, listed as synonyms marked nom. illeg.
Arachniodes amoena (Ching) Ching 1962 (Fujian, Guangdong, Guangxi, Guizhou, Hunan, Jiangxi, Zhejiang)Arachniodes arisanica (Rosenst.) Ching 1962 =
Arachniodes globisora (Hayata) Ching 1964 Arachniodes carvifolia (Kunze) Ching 1962 – East Indian holly fern (Yunnan, India, Philippines, Sri Lanka, U.S.)Arachniodes chinensis (Rosenst.) Ching 1962 Arachniodes coniifolia (T. Moore) Ching 1962 (Sichuan, Yunnan, Nepal)Arachniodes denticulata (Sw.) Ching 1962 Arachniodes diffracta (Baker) Ching 1962 =
Acrorumohra diffracta (Baker) H. Itô 1938 (Guangxi, Hainan, Yunnan, Japan, Vietnam)
Arachniodes dimorphophyllum (Hayata) Ching 1962 Arachniodes exilis (Hance) Ching 1962 Arachniodes festina (Hance) Ching 1962 (China)Arachniodes grossa (Tardieu & C.Chr.) Ching 1962 (Guangdong, Hainan)Arachniodes hasseltii (Blume) Ching 1962 =
Acrorumohra hasseltii (Blume) Ching 1964 (Hainan, Taiwan, India, Nepal)
Arachniodes hainanensis (Ching) Ching 1962 (Hainan)Arachniodes henryi (H.Christ) Ching 1962 Arachniodes leucostegioides (C.Chr.) Ching 1962 Arachniodes nigrospinosa (Ching) Ching 1962 Arachniodes obtusissima (Ching) Ching 1962 Arachniodes simulans (Mett. ex Kuhn) Ching 1962 Arachniodes speciosa (D.Don) Ching 1962 Arachniodes spectabilis (Ching) Ching 1962 Arachniodes subreflexipinna (Ogata) Ching 1962 =
Acrorumohra subreflexipinna (Ogata) H. Itô in Nakai & Honda 1938 Arachniodes tonkinensis (Ching) Ching 1962 Arachniodes zeylanica (Ching) Ching 1962
In 1962, Satoru Kurata published descriptions of the following specie in the Science Report of the Yokosuka City Museum , Yokosuka, Japan.
Arachniodes × kenzo-satakei (Sa.Kurata) Sa.Kurata 1962 Arachniodes subamabilis Sa.Kurata 1962 Ren Chang Ching and Chu Hao Wang published the following species in Acta Phytotaxonomica Sinica , volume 9, number 4, 1964.
Arachniodes acuminata Ching & Chu H.Wang 1964 =
Arachniodes sphaerosora (Tagawa) Ching 1965 Arachniodes caudata (Tagawa) Ching 1964 Arachniodes globisora (Hayata) Ching 1964 Arachniodes obtusiloba Ching & Chu H.Wang 1964 Arachniodes pseudocavalerii Ching & Chu H.Wang 1964 Arachniodes rhomboidea (Schott) Ching 1964 Arachniodes yoshinagae (Makino) Ching 1964
Ren Chang Ching in Acta Phytotaxonomica Sinica , volume 10, number 3, 1965, published:
Arachniodes sphaerosora (Tagawa) Ching 1965 Satoru Kurata published the following Arachniodes species name in the Journal of Geobotany (Hokuriku Journal of Botany) , volume 13, in 1965.
Arachniodes hekiana Sa.Kurata 1965 Published by George Richardson Proctor in the British Fern Gazette . volume 9, number 6, 1965
Arachniodes denticulata (Sw.) Proctor 1965 (nom. illeg.) =
Arachniodes denticulata (Sw.) Ching 1962 Arachniodes rigidissima (Hook.) Proctor 1965 Edmund André Charles Louis Eloi Schelpe published the following species in Boletim da Sociedade Broteriana , sér. 2, vol. 41, 1967.
Arachniodes webbiana Schelpe 1967 Arachniodes foliosa Schelpe 1967 In 1968, Satoru Kurata published the following Arachniodes species names in the Journal of Geobotany (Hokuriku Journal of Botany) , volume 16.
Arachniodes repens Sa.Kurata 1968 Arachniodes yasu-inouei Sa.Kurata 1968 Arachniodes × masakii Sa.Kurata 1968 Arachniodes × clivorum Sa.Kurata 1968 Arachniodes × kurosawae Shimura & Sa.Kurata in Sa.Kurata 1968 Arachniodes × pseudohekiana Sa.Kurata 1968 Arachniodes × sasamotoi Sa.Kurata 1968
Toshiyuki Nakaike published the following new species in Journal of Geobotany (Hokuriku Journal of Botany) , volume 17, 1969, and volume 18, 1970.
Arachniodes okinawensis Nakaike 1969 (Japan)Arachniodes pseudorepens Nakaike 1970 (Japan)In Flora Tsinlingensis , volume 2, 1974, Ren Chang Ching published:
Arachniodes centrochinensis Ching 1974 =
Arachniodes simulans (Ching) Ching 1962 (China)
Rolla Milton Tryon and Davis S. Conant published the following species in Acta Amazonica , volume 5, number 1, in 1975.
Arachniodes macrostegia (Hook.) R.M.Tryon & D.S.Conant 1775 (Brazil, Venezuela)In 1975, Toshiyuki Nakaike published the following species names in his Enumeratio Pteridophytarum Japonicarum, Filicales .
Arachniodes sporadosora (Kunze) Nakaike 1775 Arachniodes japonica (Kurata) Nakaike 1775 Satoru Kurata published the following Arachniodes species names in the Journal of Geobotany (Hokuriku Journal of Botany) , volume 25, in 1978.
Arachniodes cantilenae Sa.Kurata 1978 Arachniodes hiugana Sa.Kurata 1978 Arachniodes kenzo-satakei Sa.Kurata 1978 Arachniodes tomitae Sa.Kurata 1978 Arachniodes × minamitanii Sa.Kurata 1978 (A. hiugana × A. nipponica )Arachniodes × mirabilis Sa.Kurata 1978 (A. assamica × A. rhomboidea )Arachniodes × mitsuyoshiana Sa.Kurata 1978 (A. miqueliana × A. simplicior )Arachniodes × respiciens Sa.Kurata 1978 (A. exilis × A. simplicior )Arachniodes argillicola (Proctor) Proctor 1982 Arachniodes chaerophylloides (Poir.) Proctor 1982 – false holly fern
Yin Tang Hsieh in Acta Botanica Yunnanica , volume 5, number 1, 1983.
Arachniodes jinpingensis Y.T.Hsieh 1983 (Yunnan)Arachniodes valida Y.T.Hsieh 1983 (Yunnan)Arachniodes dayaoensis Y.T.Hsieh 1983 (Yunnan)Yin Tang Hsieh in Bulletin of Botanical Research , volume 4, number 2, 1984.
Arachniodes caudifolia Ching & Y.T.Hsieh 1984 (Yunnan)Arachniodes gansuensis (Ching) Y.T.Hsieh 1984 (Yunnan, Sichuan)Arachniodes guangnanensis Y.T.Hsieh 1984 (Yunnan)Arachniodes lushuiensis Y.T.Hsieh 1984 (Yunnan)Arachniodes reducta Y.T.Hsieh & Y.P.Wu 1984 (Zhejiang)Arachniodes subaristata Ching & Y.T.Hsieh 1984 (China)Yin Tang Hsieh and Ren Chang Ching, in Acta Phytotaxonomica Sinica , volume 22, number 2, 1984.
Arachniodes ishingensis Ching & Y.T.Hsieh 1984 Arachniodes obtusipinnula Ching & Y.T.Xie 1984 =
Arachniodes tonkinensis (Ching) Ching 1962 Arachniodes ziyunshanensis Y.T.Hsieh 1984 Ren Chang Ching and Zheng-yu Liu, in Acta Phytotaxonomica Sinica , volume 22, number 4, 1984.
Arachniodes nanchuanensis Ching & Z.Y.Liu 1984 (Sichuan)Ren Chang Ching published the following species names in the Bulletin of Botanical Research , volume 6, number 3, in 1986.
Arachniodes abrupta Ching 1986 (Guangxi, Zhejiang)Arachniodes ailaoshanensis Ching 1986 (Yunnan)Arachniodes anshunensis Ching & Y.T. Hsieh 1986 (Guizhou)Arachniodes aristatissima Ching 1986 (Zhejiang)Arachniodes attenuata Ching 1986 (Yunnan)Arachniodes austro-yunnanensis Ching 1986 (Yunnan)Arachniodes baiseensis Ching 1986 (Guangxi)Arachniodes calcarata Ching 1986 (Guizhou, Hunan, Sichuan)Arachniodes cyrtomifolia Ching 1986 (Sichuan)Arachniodes cornopteris Ching 1986 (Sichuan)Arachniodes costulisora Ching 1986 (Sichuan)Arachniodes decomposita Ching 1986 (Guizhou)Arachniodes elevatas Ching 1986 (Sichuan)Arachniodes emeiensis Ching 1986 (Sichuan)Arachniodes erythrosora Ching 1986 =
Arachniodes festina (Hance) Ching 1962 (Guangdong)
Arachniodes falcata Ching 1986 (Sichuan)Arachniodes fengii Ching 1986 Arachniodes foeniculacea Ching 1986 Arachniodes fujianensis Ching 1986 Arachniodes gigantea Ching 1986 Arachniodes gijiangensis Ching 1986 Arachniodes gizushanensis Ching 1986 Arachniodes gongshanensis Ching & Y.T.Hsieh 1986 Arachniodes gradata Ching 1986 Arachniodes guangtongensis Ching 1986 Arachniodes guangxiensis Ching 1986 Arachniodes guanxianensis Ching 1986 Arachniodes hekouensis Ching 1986 Arachniodes heyuanensis Ching 1986 Arachniodes huapingensis Ching 1986 Arachniodes hunanensis Ching 1986 Arachniodes jiangxiensis Ching 1986 Arachniodes jinfoshanensis Ching 1986 Arachniodes jingdongensis Ching 1986 Arachniodes leuconeura Ching 1986 Arachniodes liyangensis Ching 1986 Arachniodes longipinna Ching 1986 Arachniodes lushanensis Ching 1986 Arachniodes maoshanensis Ching 1986 Arachniodes mengziensis Ching 1986 Arachniodes multifida Ching 1986 Arachniodes nanqingensis Ching 1986 Arachniodes neoaristata Ching 1986 Arachniodes nitidula Ching 1986 Arachniodes pianmaensis Ching 1986 Arachniodes pseudo-assamica Ching 1986 Arachniodes pseudo-longipinna Ching 1986 Arachniodes pseudo-simplicior Ching 1986 Arachniodes semifertilis Ching 1986 Arachniodes setifera Ching 1986 Arachniodes sichuanensis Ching 1986 Arachniodes similis Ching 1986 Arachniodes sino-aristata Ching 1986 Arachniodes sino-rhomboidea Ching 1986 Arachniodes shuangbaiensis Ching 1986 Arachniodes sparsa Ching 1986 Arachniodes spino-serrulata Ching 1986 Arachniodes subamoena Ching 1986 =
Arachniodes amoena (Ching) Ching 1962 Arachniodes triangularis Ching 1986 Arachniodes xinpingensis Ching 1986 Arachniodes yaomashanensis Ching 1986 Arachniodes yinjiangensis Ching 1986 Arachniodes yunnanensis Ching 1986 Arachniodes borealis Seriz. 1986 (Japan, Korea)=
Leptorumohra borealis ( Seriz. ) Shmakov 2009
Yin Tang Hsieh published the following species names in the Bulletin of Botanical Research , volume 6, number 4, in 1986.
Arachniodes chingii Y.T.Hsieh 1986 Arachniodes damiaoshanensis Y.T.Hsieh 1986 Arachniodes futeshanensis Y.T.Hsieh 1986 Arachniodes nibashanensis Y.T.Hsieh 1986 Arachniodes yunqiensis Y.T.Hsieh 1986 Ren Chan Ching and Yin Tang Hsieh, in the Bulletin of Botanical Research volume 11, number 2, 1991, published:
Arachniodes basipinnata (Ching) Ching ex Y.T.Hsieh 1991 Arachniodes lanceolata Y.T.Hsieh 1991 Arachniodes menglianensis Y.T.Hsieh 1991 Arachniodes parasimplicior Ching ex Y.T.Hsieh 1991 =
Arachniodes simplicior (Makino) Ohwi 1962 Arachniodes fengyangshanensis Ching & C.F.Zhang ex Y.T.Hsieh 1991 =
Arachniodes exilis (Hance) Ching 1962 Arachniodes davalliaeformis (Christ) Nakaike 1992 – shiny bristle fernArachniodes insularis W.H.Wagner 1993 – Santa Cruz Island holly fernThe following two species were published by Shi Fu Wu in Keys to the Vascular Plants of the Wuling Mountains Area (edited by Wang Wentsai) 572–573, pl. 5. Beijing 1995.
Arachniodes hupingshanensis S.F.Wu 1995 (China)Arachniodes wulingshanensis S.F.Wu 1995 (China)Arachniodes squamulosa R.C.Moran & B.Øllg. 1998 In 2008, Christopher Roy Fraser-Jenkins published the following new Arachniodes species in his Taxonomic revision of three hundred Indian subcontinental pteridophytes with a revised census-list: a new picture of fern-taxonomy and nomenclature in the Indian subcontinent .
Arachniodes cornucervi (D.Don) Fraser-Jenk. 2008 Arachniodes palmipes (Kunze) Fraser-Jenk. 2008 Arachniodes sledgei Fraser-Jenk. 2008 The genus Leptorumohra was segregated from the genus Rumohra in 1938 by Hiroshi Itô. In 1962, Jisaburo Ohwi included at least three of the Leptorumohra species in Arachniodes , but these placements were not universally accepted. In 2006, Alan Reid Smith & al. in "A classification for extant ferns" included the genus Leptorumohra in Arachniodes . Leptorumohra species, which may be considered part of the Arachniodes genus include:
Leptorumohra amurensis (H.Christ) Tzvelev 1989 (Siberia, Poland, Japan, Heilongjiang, Jilin, Liaoning, Mongolia)Arachniodes borealis Seriz. 1986 (=)
Leptorumohra borealis ( Seriz.) Shmakov 2009 Arachniodes miqueliana (Maxim. ex Franch. & Sav.) Ohwi 1962 (=)
Polystichopsis miqueliana (Maxim. ex Franch. & Sav.) Tagawa 1958 (=)
Leptorumohra miqueliana (Maxim. ex Franch. & Sav.) H. Itô 1938 (=)
Rumohra miqueliana (Maxim. ex Franch. & Sav.) Ching 1934 (=)
Dryopteris miqueliana (Maxim. ex Franch. & Sav.) C.Chr 1905 (=)
Nephrodium miquelianum (Maxim. ex Franch. & Sav.) Kom. 1901 (=)
Aspidium miquelianum Maxim. ex Franch. & Sav. 1876 Arachniodes mutica (Franch. & Sav.) Ohwi 1962 (=)
Polystichopsis mutica (Franch. & Sav.) Tagawa 1958 (=)
Leptorumohra mutica (Franch. & Sav.) Czerep. 1981 (=)
Rumohra mutica (Franch. & Sav.) Ching 1934 (=)
Dryopteris mutica (Franch. & Sav.) C.Chr. 1905 (=)
Aspidium muticum Franch. & Sav. 1876 Arachniodes quadripinnata (Hayata) Seriz. 1986 (=)
Leptorumohra quadripinnata (Hayata) H.Itô 1938 (=)
Microlepia quadripinnata Hayata 1911 Arachniodes sinomiqueliana (Ching) Ohwi 1962 (=)
Polystichopsis sinomiqueliana (Ching) Tagawa 1958 (=)
Leptorumohra sinomiqueliana (Ching) Tagawa 1939 (=)
Rumohra sinomiqueliana Ching 1934